Page is the fourth president and director of the Whitehead Institute for Biomedical Research, a professor of biology at MIT, and an Investigator of the Howard Hughes Medical Institute. A graduate of Swarthmore College, Page earned his M.D. from Harvard Medical School and the Harvard-MIT Health Sciences and Technology program. He joined the Whitehead Institute as the first Whitehead Fellow in 1984. Page’s honors include a MacArthur Prize Fellowship, Science magazine’s Top Ten Scientific Advances of the Year (1992 and 2003), the Francis Amory Prize of the American Academy of Arts and Sciences, and the March of Dimes Prize in Developmental Biology. He is a member of the National Academy of Sciences, the National Academy of Medicine, and the American Academy of Arts and Sciences.